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Recent nonculture-based efforts that prepare and sequence clone libraries (see Section 67) from sampled human feces, ocean water, and sampled marine sediments (Edwards and Rohwer, 2005) have provided evidence for ~1000 viral genotypes in human feces, ~5000/200 L in seawater, and up to 106 in marine sediment The vast majority of these environmental virus sequences differ immensely from those of domesticated viruses Lytic viruses threaten to obliterate prokaryotic populations, whereas lysogenic and chronic infections represent parasitic interactions Virus-induced mortality of prokaryotes is highly variable in time and space dependent especially on habitat characteristics, host abundances, and the host s physiological state Because viral infections are host-speci c and density-dependent, it is widely accepted that infections can control the composition of prokaryotic communities Competitively dominant prokaryotes can be kept in check because viruses kill the winner (Weinhauer, 2004).

Regarding biogeochemistry, viruses are in uential via at least two mechanisms: (i) lytic cycles can reduce the sizes, hence physiological activity of prokaryotes that catalyze individual steps in the cycling of carbon, nitrogen, sulfur, and other elements; and (ii) via the viral shunt , nutrients can be moved from the cell-associated nutrient pool to the particulate and dissolved nutrient pool Figure 520 illustrates the viral shunt for an oceanic habitat In the classic view of trophic dynamics, microbial phytoplankton are grazed by zooplankton, which serve as food for carnivores If lytic viruses divert phytoplankton biomass into the pool of dissolved and particulate organic matter, there are two potential conseFigure 520 Viral in uences on biogeochemistry Viruses cause the lysis of cells, converting them into quences: the organic matter is utilized as particulate organic matter (POM) and dissolved organic a carbon source by heterotrophic bacmatter (DOM).

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FIGURE 15.5 The Language pop-up menu on the Character panel lets you choose a different language for hyphenating and spell-checking highlighted text.

This reduces the rate at which carbon teria (producing CO2) and the ux of sinks from the surface layer into the deep ocean where particular carbon (detritus) reaching the carbon is trapped for millennia (biological carbon and stored in marine sediments is pump) Instead the carbon is retained in the surface diminished (Suttle, 2005) Both of these waters where it is photooxidized and respired, in consequences may contribute to global chemical equilibrium with the atmosphere The net warming (see Section 72) For addieffect is a faster rate of CO2 buildup in the atmosphere tional information on viruses, see Granthan would occur if the POM were exported to the off and Webster (2005), Cann (2005), deep ocean (Reprinted by permission from Macmillan Villarreal (2005), Knipe et al (2001), and Publishers Ltd: Nature, from Suttle, CA 2005 Viruses in the sea Nature 437:356 361 Copyright 2005) Hurst (2000)..

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If no document is open when you create, edit, or delete colors, the new color palette becomes the default for all future documents. n

Figure 11-1: This chart shows the best-fit line for a set of data points showing the relationship between customers tenures and the amount they have paid.

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5 In order for users on two systems to communicate, a common set of rules must be implemented. The International Standards Organization recognized the need in 1977 for the standardization of information-network architectures. ISO s TC97 committee, responsible for information systems, created the Open Systems Interconnection (OSI) model in 1979. In uenced by the earlier network architectures of IBM (SNA 1974) Systems Network Architecture: Technical Overview (Third Edition), IBM Corporation, September 1986 and the Digital Equipment Corporation (DNA 1975) DECnet DIGITAL Network Architecture (Phase V): General Description , Digital Equipment Corporation, September 1987, the ISO Open System Interconnection model divided the services needed to implement computer communication into layers.

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