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Some environmentally induced traits persist for several generations For example, a particular Drosophila strain that normally grows at 21 C was exposed to 36 C for twenty-two hours Dwarf progeny were produced When they were mated among themselves, fewer and fewer dwarfs appeared in each generation, but smallerthan-normal ies were produced as late as the fth generation The appearance of an environmentally induced trait that persists for several generations has been termed dauermodi cation Extrachromosomal inheritance is usually identi ed by the odd results of reciprocal crosses If the progeny of reciprocal crosses are not followed for several generations, the results can be misleading when extrachromosomal inheritance is involved Where feasible, nuclear transplantation has proved useful in identifying extrachromosomal inheritance In this technique, the nucleus of a cell, such as an amoeba or frog egg, is removed by microsurgery or destroyed by radiation, and another nucleus substituted.

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layer of carbon onto a nonconducting cylindrical substrate (see Figure P224) If such a cylinder has

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Thus, not only can a nucleus be isolated from its cytoplasm, but various nuclei can be implanted in the same cytoplasm A similar experiment, called a heterokaryon test, can be done with various fungi such as Neurospora and Aspergillus: Mycelia can fuse and form a heterokaryon, a cell containing nuclei from different strains Thus, nuclei of both strains exist in the mixed cytoplasm Subsequently, spores (conidia) that have one or the other nucleus in the mixed cytoplasm can be isolated The phenotype of the colonies produced from these isolated conidia show whether the trait under observation is controlled by the nucleus or the cytoplasm Chromosomal genes in a particular cytoplasm can also be isolated by repeated backcrossing of offspring with the male-parent type In each cross, the content of the female chromosomal genes is halved, but, presumably, the cytoplasm remains similar to the female line.

(Contd )

2

Thus, after several generations, male genes can be isolated in female cytoplasm The phenotypic results of the nal cross will indicate whether inheritance was chromosomal or extrachromosomal..

radius a and length d, determine the thickness of the lm required for a resistance R if: a = 1 mm S 1 = = 29 M m R = 33 k d = 9 mm

however, may not be exactly the same; that is, their tolerances are such that the resistances may not be exactly 10 k a If the resistors have 10 percent tolerance, nd the worst-case output voltages b Find these voltages for tolerances of 5 percent

Snails are coiled either to the right (dextrally) or to the left (sinistrally) as determined by holding the snail with the apex up and looking at the opening. The snail is dextrally coiled if the opening comes from the right-hand side and sinistrally coiled if it comes from the left-hand side ( g. 17.1).The inheritance pattern of the coiling is at rst perplexing.

Neglect the end surfaces of the cylinder and assume that the thickness is much smaller than the radius

Schlumberger A dual Hydrashaker system providing 2 DoF used to test oil well down-hole electronic logging/drilling tools The double pivot Hydrostatic Spherical Couplings allow for angular misalignment while providing a backlash free, direct load path between the Hydrashakers and the shake table

resistor values (including the power rating) necessary to achieve the indicated voltages 1 Resistors are available in 1 -, 1 -, 2 -, and 1-W ratings 8 4

The McGraw Hill Companies, 2001

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Figure 17.1 Inheritance of coiling in the pond snail, Limnaea peregra. Reciprocal crosses (D, dominant dextral; d, recessive sinistral coiling) are shown (DD mated with dd in each case). The F1 individuals in both crosses have the Dd genotype but re ect the mother s genotype in respect to coiling; DD mothers produce dextrally coiled offspring, whereas dd mothers produce sinistrally coiled offspring. The F2 individuals in both cases are identical because the genotypes of the F1 mothers are identical (Dd ). The coiling of a snail s shell is determined by its mother s genotype, not phenotype.

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